Oscar (Will Smith), a lowly tonguescrubber at the local Whale Wash, becomes a hero when he tells a great white lie. When his lie begins to unravel, Oscar teams up with an outcast vegetarian shark, Lenny (Jack Black) and his loyal friend Angie (Renée Zellweger) to help him stand up to the most feared shark in the water (Robert De Niro) and find his true place in the reef.