In this video Paul Andersen explains how the frequency of recombination between linked genes can be used to determine the relative location of genes on a chromosome. Thomas Hunt Morgan and Alfred Strutevant used the fruit fly to develop a theory of chromosomal inheritance and discover crossing over.

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Music Attribution
Title: String Theory
Artist: Herman Jolly
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"File:Drosophila Repleta Lateral.jpg." Wikipedia, the Free Encyclopedia. Accessed March 13, 2014. http://en.wikipedia.org/wiki/File:Dro....
"File:Morgan Crossover 1.jpg." Wikipedia, the Free Encyclopedia. Accessed March 13, 2014. http://en.wikipedia.org/wiki/File:Mor....
"File:Thomas Hunt Morgan.jpg." Wikipedia, the Free Encyclopedia. Accessed March 13, 2014. http://en.wikipedia.org/wiki/File:Tho....
"FlyBase," n.d. http://flybase.org/reports/FBgn000397....
spax89. Illustration of a Tobacco Pipe, 2009. Extracted from Media:Blason de la ville de SaintQuentinlaPoterie (30).svg. http://commons.wikimedia.org/wiki/Fil....