During the Early Paleocene, Placental mammals began to diversify explosively after the extinction of the NonAvian dinosaurs. This included the ancestors of the hoofed ungulates, which often looked almost nothing like their familiar modern relatives. Indeed, many were very generalized animals, more closely resembling Carnivorans, Hyraxes and guinea pigs. Basal were omnivorous and sometimes even carnivorous, although adaptations for herbivory and cursorial behavior developed multiple times as well.

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